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From The Collaborative International Dictionary of English v.0.48:
Vaginant \Vag"i*nant\, a. [Cf. F. vaginant. See Vagina.] Serving to invest, or sheathe; sheathing. [1913 Webster] Vaginant leaf (Bot.), a leaf investing the stem or branch by its base, which has the form of a tube. [1913 Webster] Vaginate